Players control an anonymous hero out to stop the evil Piraka, who have used the power of the Mask of Life to transform the wildlife of the island of Voya Nui into aggressive, evil creatures. The hero is able to wear the masks of the Toa Inika in order to take on their appearances and powers. Players can switch between any collected masks at will, but if they run out of health the current mask is lost and a new one must be found before the character can become that Toa again. Collecting a mask also restores health, even if it is one the character already has.
During gameplay, players collect Lego pieces with different point values. Collecting enough points fills a meter and activates "Hero Mode", turning the character golden and giving invincibility as well as an offensive boost. After completing a level, points collected can also be used to purchase items and upgrades from the in-game Lego Shop. Players are able to upgrade weapons throughout the game, starting with the Toa Mata's tools before upgrading to the Toa Metru's tools and finally the Toa Inika's tools; armor and special abilities can also be upgraded.
Throughout each level are several Lego constructions, or construction-actions, that can be assembled. Two kinds that must be activated to proceed are normal constructions activated by Toa Hewkii and golden constructions that can be activated by anyone in Hero Mode, though doing so causes Hero Mode to end and more points must be collected before it can be entered again. Also found are character-specific constructions that provide bonuses when activated.
Scattered throughout the levels are several canisters that give bonuses when collected: collecting all of the silver canisters in a level unlocks a common enemy for viewing between levels, while each gold canister found unlocks a collectible item for viewing. Each regular level includes five silver canisters and four gold canisters, and Piraka levels include four gold canisters apiece. Boss enemies-which include the seven Piraka, the six Rahkshi from the 2003 story, and the large set characters Nidhiki, Krekka, Roodaka, Sidorak, Brutaka, and Axonn-are also unlocked for viewing after they have been defeated.
There are 25 levels and 19 bosses in total. In addition to the levels and viewable collectibles, players can also explore a bonus area called Piraka Playground, where the Piraka can be seen walking around after you defeat them in the main levels. Players can purchase additional items for Piraka Playground from the Shop and watch the Piraka use them. There are also 3 bonus levels available for purchase, which are accessed through Piraka Playground.
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