The Fire Totem in Skylanders refers to the Fire Totem Trap Crystal, a collectible from Skylanders Trap Team, also known as the Searing Spinner, used to capture Fire villains and utilize their powers in the game. It's a Traptanium crystal that harnesses the Fire element, fitting into the Portal of Power alongside Skylanders figures to enhance gameplay, especially in turret challenges where fire-based attacks are key.